Beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died at about 141, the zoo says, adding that ...

No one knows exactly when Gramma was born. But if the estimated birth year of 1884 is accurate, Chester Arthur occupied the ...

Gramma the Galápagos tortoise died on Nov. 20, 2025. The oldest animal at the San Diego Zoo was believed to be around 141 ...

Gramma, who died at the zoo on Thursday, was believed to have been born in 1884, when Chester A. Arthur was in the White ...

The San Diego Zoo's oldest resident, a Galapagos tortoise named Gramma, died Nov. 20. She was estimated to be 141 years old.

M ore than a dozen islands, formed millions of years ago, rise out of the Pacific Ocean 1,000km (600 miles) from Ecuador’s ...

Gramma, a Galápagos tortoise described by her care specialists as “the queen of the zoo,” was euthanized last Thursday after ...

The San Diego Zoo has said goodbye to one of its most cherished icons, Gramma, a Galápagos tortoise who inspired generations ...

SAN DIEGO, California — Gramma, a Galápagos tortoise and the oldest known resident of the San Diego Zoo, has died at an ...

One of the most comprehensive surveys to date of shark and other large predator fish in the Eastern Tropical Pacific (ETP) ...

Gramma the tortoise, believed to be 141 years old and the oldest resident of the San Diego Zoo, was euthanized after ...

Gramma, an estimated 141-year-old Galápagos tortoise, died Thursday at the San Diego Zoo, ending a near century-long chapter ...