Gramma the Galápagos tortoise, San Diego Zoo's oldest resident, dies at about 141

Beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died at about 141, the zoo says, adding that ...

Meet Gramma: The 141-year-old Galápagos tortoise died who witnessed two world wars died at the San Diego Zoo

San Diego Zoo mourns the loss of Gramma, a Galápagos tortoise estimated to be 141 years old. Born in the Galápagos Islands, ...

How a 'sweet and shy' tortoise outlived empires and survived two world wars

No one knows exactly when Gramma was born. But if the estimated birth year of 1884 is accurate, Chester Arthur occupied the ...

Gramma, a Galápagos Tortoise Older Than the San Diego Zoo, Dies at About 141

Gramma, who died at the zoo on Thursday, was believed to have been born in 1884, when Chester A. Arthur was in the White ...

'Queen' Gramma the Tortoise, the Oldest Animal at the San Diego Zoo, Dies at the Estimated Age of 141

Gramma the Galápagos tortoise died on Nov. 20, 2025. The oldest animal at the San Diego Zoo was believed to be around 141 ...

Zoo mourns giant tortoise that lived over 140 years

The San Diego Zoo's oldest resident, a Galapagos tortoise named Gramma, died Nov. 20. She was estimated to be 141 years old.

'Queen of the zoo' dies: Gramma the Galapagos tortoise was about 141 (Associated Press video)

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise ...
The Economist

How to save the Galápagos from its visitors

M ore than a dozen islands, formed millions of years ago, rise out of the Pacific Ocean 1,000km (600 miles) from Ecuador’s ...

A San Diego Zoo Galápagos Tortoise, Thought to Be 141, Is Dead

Gramma, a Galápagos tortoise described by her care specialists as “the queen of the zoo,” was euthanized last Thursday after ...

140-Year-Old 'Gramma', The Galapagos Tortoise Passes Away In San Diego Zoo

The San Diego Zoo has said goodbye to one of its most cherished icons, Gramma, a Galápagos tortoise who inspired generations ...

Shark strongholds: Remote Pacific islands host thriving populations as coastal marine reserves falter

One of the most comprehensive surveys to date of shark and other large predator fish in the Eastern Tropical Pacific (ETP) ...

Gramma the Galápagos tortoise, an original San Diego Zoo resident, dies at 141

Gramma the tortoise, believed to be 141 years old and the oldest resident of the San Diego Zoo, was euthanized after suffering from progressive bone deterioration because of her age.